New calcareous dinoflagellates (Calciodinelloideae) from the Middle Coniacian to Upper Santonian chalks of Lägerdorf (northern Germany)

Abstract. Three new calcareous dinoflagellate species from the Middle Coniacian to Upper Santonian chalks of Lägerdorf (northern Germany) are formally described: Calcicarpinum macrogranulum n. sp., Pirumella fragilis n. sp. and Ruegenia quinqueangulata n. sp. The species show differing vertical distribution patterns which might result from local sea-level changes: P. fragilis and R. quinqueangulata are restricted to the possibly transgressive upper Mid-Coniacian to Lower Santonian interval and C. macrogranulum occurs consistently only in the probably regressive lower Mid-Coniacian and Middle to Upper Santonian intervals.