Articles | Volume 37, issue 1
Research article
16 May 2018
Research article |  | 16 May 2018

New species of Mesozoic benthic foraminifera from the former British Petroleum micropalaeontology collection

Lyndsey R. Fox, Stephen Stukins, Tom Hill, and Haydon W. Bailey

This paper describes five new Mesozoic, deep-water benthic foraminifera from the former British Petroleum microfossil reference collections at the Natural History Museum, London. The focus is on selected calcareous and agglutinating taxa that are of stratigraphical and/or palaeoecological significance for academic and industrial related activities.

Ophthalmidium dracomaris (, Trochammina fordonensis (, Eobigenerina calloviensis (, Arenoturrispirillina swiecickii ( and Ataxophragmium mariae ( are described with new illustrations. Their biostratigraphic and palaeoecological significance are briefly discussed.

1 Introduction

The North Sea Basin lies in between Scandinavia and north-western Europe and is bounded by the continental shelf edge, approximately comparable to the region covered by the current North Sea today. It is a topographical low and can be divided into several sub-basins of late Palaeozoic through Cenozoic age (Carr, 2004). The area has an active tectonic history and deposition has varied between the different basins. Benthic foraminifera have been used extensively in the oil and gas industry to date the sediments and investigate the impact of environmental and palaeobathymetric changes on these semi-enclosed basins (Gradstein et al., 1994; Jenkins and Murray, 1989; King and Hughes, 1983).

British Petroleum's exploration activities, especially in the UK sector, during the middle of the 20th century amassed a large volume of Mesozoic sample material and subsequently prepared micropalaeontological specimens, many of which are now housed at the Natural History Museum, London. The north-western European reference collection contains many of the specimens used in exploration activities and provides an almost unique collection of useful, yet in some instances formally un-described, foraminiferal taxa.

The taxonomy of the Cenozoic material from this collection has already been the focus of separate study (Fox et al., 2018). Presented here are five newly named species of Mesozoic benthic foraminifera which have academic and industrial value in their biostratigraphic and palaeoecological significance.

2 Materials

Light microscope images were taken using a Nikon SMZ25 stereoscopic zoom microscope and Nikon DS-Fi2 digital camera. SEM images were taken using uncoated specimens on a LEO 1455 VP scanning electron microscope.

3 Systematics

We present systematic data (descriptions, synonymies, biogeography and palaeoecologies) that were assembled from primary research, from the micropalaeontological literature, and also from consultation with specialists. Brief geological context is also provided, where available, for the well and/or outcrop where the type specimens were found. The taxonomic classification follows Loeblich and Tappan (1987) with a few exceptions which mainly reflect taxonomic updates published subsequently (Kaminski, 2014).

Order Miliolida Delage & Hérouard, 1896

Superfamily Nubecularioidea Jones, 1875

Family Ophthalmidiidae Wiesner, 1920

Genus Ophthalmidium Kubler & Zwingli, 1870

Type species: Oculina porosa Kubler & Zwingli, 1866

Ophthalmidium dracomaris sp. nov.

Figs. 1.1–1.5


Ophthalmidium sp. A Coleman in Horton et al. 1974.

Ophthalmidium sp. A Medd in Richardson 1979.

Ophthalmidium sp. A Morris & Coleman 1981, 221. pl. 6.3.6, fig. 15.

Diagnosis: Compressed porcellaneous test, consisting of six to eight chambers, with a long tapering neck and simple circular aperture.

Dimensions: Diameter is  100 µm; height of test is  300 µm.

Description: Test compressed, elliptical in outline; porcellaneous wall texture, periphery acute and weakly keeled; chambers almost invisible externally, half-whorl in length, rounded at the base and tapering slightly towards the aperture, margins almost parallel, sutures very weakly depressed; aperture is simple, circular, at the end of a long tapering neck, lacking a tooth.

Remarks: Specimens were originally described by Coleman (1974; in Hortman et al., 1974) from the area around Milton Keynes, Buckinghamshire, and these have been described in some detail by Morris and Coleman (1989). Ophthalmidium dracomaris sp. nov. differs from O. compressum (Barnard, Cordey & Shipp, 1981) by possessing fewer chambers. O. maredraco bears a resemblance to O. michalskii illustrated in Plate 1, Figs. 1, 6 and 7, but differs by having a more flattened, less rounded test and a longer, narrower neck.

Type level: Middle Jurassic. Late Callovian, athleta–lamberti ammonite zones.

Type locality: Well: Tetney Lock-1, Lincolnshire, UK. Depth: 1652 ft (503.5 m).

Distribution: Late Callovian of England.

Etymology: This species was named based upon its resemblance to the head of a sea dragon.

Holotype: NHMUK PM PF 74496.

Paratypes: NHMUK PM PF 74497–74500.

Plate 1(1–5) Ophthalmidium dracomaris n. sp.: (1) holotype (NHMUK PM PF 74496); (2–5) paratypes (NHMUK PM PF 74497–74500). (8–9) Eobigenerina calloviensis n. sp.: (8) holotype (NHMUK PM PF 74504); (9) paratype (NHMUK PM PF 74505). (10–12) Trochammina fordonensis n. sp.: (10) holotype (NHMUK PM PF 74501); (11–12) paratype (NHMUK PM PF 74502–74503). (13–15) Arenoturrispirillina swiecickii n. sp.: (13) holotype (NHMUK PM PF 74506); (14–15) paratype (NHMUK PM PF 74507 + NHMUK PM PF 74552). (16–17) Ataxophragmium mariae n. sp.: (16) holotype (NHMUK PM PF 74553); (17) paratype (NHMUK PM PF 74554).


Order Lituolida Lankester, 1885

Superfamily Trochamminoidea Schwager, 1877

Family Trochamminidae Schwager, 1877

Genus Trochammina Parker & Jones, 1859

Type species: Rotalia (Trochammina) inflata (Montagu) =Nautilus inflatus Montagu, 1808

Trochammina fordonensis sp. nov.

Figs. 1.10–1.12


Trochammina sp. 68 Internal British Petroleum report.

Diagnosis: Small, low spired, finely agglutinated, trochoid test with eight to nine chambers in the final whorl, umbilical to extra-umbilical aperture with a lip.

Dimensions: Diameter is 150–200 µm; height of test is  200 µm.

Description: Test small, trochoid, flattened, very low spired, rounded in outline; consists of two whorls with eight to nine chambers in the final whorl, increasing gradually in size, flat on umbilical surface, convex on spiral side. Periphery acute, slightly lobate on margin. On the umbilical side only the final whorl is visible. Sutures are radial on umbilical side and slightly curved on the spiral side. Wall finely agglutinated with much cement; umbilical to extra-umbilical aperture with a lip, opening into umbilicus commonly obscured due to preservation.

Remarks: Trochammina fordonensis sp. nov. is similar to Trochammina aquilonaris Hedinger, 1993 in general morphology and size, but differs in having a flattened dorsal side and radial sutures on the umbilical side.

Type level: Early Aptian.

Type locality: Well: Fordon-1 (BGS Borehole TA07NE24), Yorkshire, UK. Depth: 750 ft (228.6 m).

Distribution: Speeton Clay Formation, UK.

Etymology: This taxon is named after the type locality.

Holotype: NHMUK PM PF 74501.

Paratypes: NHMUK PM PF 74502–74503.

Order Textulariida Delage & Hérouard, 1896

Suborder Verneuilinina Mikhalevich & Kaminski, 2004.

Superfamily Verneuilinoidea Cushman, 1911.

Family Prolixoplectidae Loeblich & Tappan, 1985.

Genus Eobigenerina Cetean, Setoyama, Kaminski, Neagu, Bubík, Filipescu & Tyszka, 2008

Type species: Bigenerina variablis Vašíček, 1947

Eobigenerina calloviensis sp. nov.

Figs. 1.8–1.9


Bigenerina sp. 24 Internal British Petroleum report.

Diagnosis: Small biserial to uniserial agglutinating test with six to seven slowly enlarging chambers.

Dimensions: Diameter is  150 µm; height of test is  300 µm.

Description: Test free, small, elongate in lateral outline and compressed; tapered at the base and rounded at the apertural end. Consists of two sections; initially biserial with six to seven slowly enlarging chambers; final two chambers are loosely biserial to lax-uniserial, not symmetrical, forming an uneven row with oblique sutures. Chambers in the uniserial part increase very little in size as added. Sutures are thin and depressed. Aperture terminal, in the form of a rounded opening on the slightly upwardly prolonged end of the last chamber. Wall finely agglutinated with grains less than 20 µm in size.

Remarks: The genus Eobigenerina differs from Bigenerina in possessing organic cement that is silicified in fossil specimens. The Cenozoic genus Bigenerina possesses calcareous cement with pseudopores. Many of the late Paleozoic to Mesozoic species that were originally described as Bigenerina likely belong in Eobigenerina (Cetean et al., 2011). The chamber arrangement in the new species is initially biserial, becoming loosely biserial to lax-uniserial (sensu Kaminski et al., 2011) in the final chambers. This species differs from Bigenerina elongata (Gauger, 1953) and B. ciscoensis (Cushman and Waters, 1928) in possessing a shorter uniserial part and smaller test.

Type level: Early Jurassic; late Callovian, athleta–lamberti ammonite zones.

Type locality: Well: Tetney Lock-1 (BGS Borehole: TA30SW93), Lincolnshire, UK. Depth 1751 ft (533.7 m).

Distribution: Currently restricted to the immediate area around the type locality.

Etymology: This simple agglutinating form is named for the type level at which it is found.

Holotype: NHMUK PM PF 74504.

Paratypes: NHMUK PM PF 74505.

Order Spirillinida Hohenegger & Piller, 1975

Superfamily Ammodiscoidea Reuss, 1862

Family Ammodiscidae Reuss, 1862

Genus Arenoturrispirillina Tairov, 1956

Type species: Spirillina limbata Brady, 1879

Arenoturrispirillina swiecickii sp. nov.

Figs. 1.13–1.15


Arenoturrispirillina sp. A Swiecicki, 1980; pl. 1, figs. 3, 4.

Diagnosis: Single sub-rectangular to rectangular tubular chamber, coiling in a regular trochospire, with a circular outline.

Dimensions: Diameter is  600 µm.

Description: Test free, large; low conical spire, consisting of proloculus followed by single tubular chamber coiling in a low regular trochospire, outline circular, peripheral margin sub-rectangular to rectangular, occasionally developing angular keels; chamber size increasing gradually and uniformly, forming five to six whorls; spiral suture distinct and depressed; aperture arcuate opening at end of chamber; wall finely agglutinated, organic cement; surface finely roughened.

Remarks: This species was referred to as Spirillina limbata Brady by Heron-Allen and Earland (1910). However, examination of their hypotypes by Barr (1962) revealed the present species to be distinct, bearing only a superficial resemblance to the Recent S. limbata. Barr (1962) described this form as “Ammodiscoides heronalleni”, but this name has never been published. This species was illustrated and described in Swiecicki's 1980 doctoral thesis in which it was removed from the genus Ammodiscoides Cushman (1909) as “it does not show the change from trochospiral to planispiral growth, characteristic of this genus” and was instead placed in Arenoturrispirillina Tairov (1956) on the basis of its “low, regular trochospiral coil”. There have been several references (e.g. Cushman, 1946; Hofker, 1959) to Cretaceous occurrences of Ammodiscoides turbinatus (Cushman, 1909) originally described from the Holocene, Gulf of Mexico. This latter species, the genotype of Ammodiscoides, “lacks the regular trochospiral form” of the present species and therefore further investigation is required on these Cretaceous references (Sweicicki, 1980).

Type level: Campanian, benthic foraminifera biozones B1ii–B5i.

Type locality: On shore, Ignaberga, Sweden.

Distribution: Rare specimens have been found in the Campanian of the Isle of Wight. A single specimen was also found in the lower Maastrichtian of Norfolk (Swiecicki, 1980).

Etymology: This species is named for Anthony Swiecicki, who first described this taxon in his doctoral thesis on the biostratigraphy of the Campanian and Maastrichtian chalks of the United Kingdom (1980).

Holotype: NHMUK PM PF 74506.

Paratypes: NHMUK PM PF 74507 + NHMUK PM PF 74552.

Order Loftusiida Kaminski & Mikhalevich, 2003

Superfamily Ataxophragmioidea Schwager, 1877

Family Ataxophragmiidae Schwager, 1877

Genus Ataxophragmium Reuss, 1860

Type species: Bulimina variablis d'Orbigny, 1840

Ataxophragmium mariae sp. nov.

Figs. 1.16–1.17


1941 Ataxogyroidina globosa (von Hagenow); Marie, p. 59, pl. 5, figs. 50-7.

1966 Ataxophragmium globosum (von Hagenow); Hofker, p. 33, p1.4, fig. 9.

1977 Ataxophragmium globosum (von Hagenow); Villain, p. 43, p1.4, figs. 2, 3.

Diagnosis: A species of Ataxophramium with a regular, low trochospiral chamber arrangement, tendency to planoconvex form, sub-rectangular, interiomarginal aperture and smooth surface.

Dimensions: Height of test: 700 µm; diameter: 650 µm.

Description: Test free; an asymmetrical low trochospiral coil with a tendency to planoconvexity, outline sub-spherical; two, occasionally three, whorls embracing, overlapping, evolute; chambers indistinct, moderately and uniformly increasing in size, may be slightly inflated, six per whorl several times higher than broad; last chamber often overlaps previous whorl both dorsally and ventrally; sutures distinct, flush; apertural face flat, making angle of approximately 65 with previous whorl; aperture a deep, elongate, sub-rectangular slit along interiomarginal suture of last chamber, may show partial development of central lobe; interior most often simple, occasional forms show development of sutural buttresses; wall finely agglutinated, with calcareous cement; surface smooth.

Remarks: Marie (1941) originally described this rather distinctive species of Ataxophragmium, noting in particular its characteristic form and aperture. However, he referred to it as Nonionina globosa von Hagenow (1842), a form described, though never figured, from the Upper Cretaceous. Later workers, including Reuss (1862), Marsson (1878), Franke (1925, 1927), Cushman (1931), Schijfsma (1946) and Visser (1951), have all considered N. globosa to be calcareous and perforate, and it is herein considered to belong to the genus Gyroidinoides. The present form A. mariae is broadly similar in shape, but its wall character and aperture serve clearly to distinguish it. Schijfsma (1946) noted the above situation and erected Ataxogyroidina pseudoglobosa and included forms referred to by Marie as A. globosa in its synonymy. From his figures and descriptions, however, it is clear that A. pseudoglobosa (Schijfsma, 1946) is not conspecific with Marie's species nor with the (Tairov, 1959) forms herein described, lacking as it does the characteristic form, aperture and smooth surface.

Type level: Upper Campanian.

Type locality: German Jura.

Distribution: This species is moderately common in the upper Campanian on the Isle of Wight and Norfolk, UK.

Etymology: Named in honour of Pierre Marie who first described this taxon.

Holotype: NHMUK PM PF 74553.

Paratypes: NHMUK PM PF 74554.

4 Summary

Five new species of deep-water benthic foraminifera from the Mesozoic of north-western Europe are described from the extensive reference collections at the Natural History Museum, London. These benthic foraminifera are important for their biostratigraphic and palaeoenvironmental value, hence their overdue formalized taxonomic descriptions. This study highlights the importance of both museum and industrial collections for their vast potential for new, publishable data, including new taxa and material from restricted and no-longer accessible locations.

Data availability

Type and figured material is deposited at the Natural History Museum, London, where it available for inspection upon request.

Competing interests

The authors declare that they have no conflict of interest.


We are grateful to our reviewers, Mike Kaminski and Nigel Ainsworth, whose constructive comments greatly improved this paper. This research was undertaken at the Natural History Museum, London, and has been funded by BP exploration (Sunbury, UK) through the project “A Re-appraisal of the BP Micropalaeontology Collection”.

TMS Promoted Paper 2018.

Edited by: Kirsty Edgar
Reviewed by: Nigel Ainsworth and Michael Kaminski


Barr, F. T.: Upper Cretaceous planktonic foraminifera from the Isle of Wight, England, Palaeontology, 4, 552–580, 1962. 

Brady, H. B.: Memoirs: Notes on some of the Reticularian Rhizopoda of the “Challenger” Expedition, J. Cell Sci., 2, 20–63, 1879. 

Carr, S. J.: The North Sea Basin, in: Quaternary Glaciations – Extent and Chronology, Part 1, edited by: Ehlers, J. and Gibbard, P. L., Elsevier, Ansterdam, 47–82, 2004. 

Cetean, C. G., Setoyama, E., Kaminski, M. A., Neagu, T., Bubík, M., Filipescu, S., and Tyszka, J.: Eobigenerina, a cosmopolitan deep-water agglutinated foraminifer, and remarks on late Paleozoic to Mesozoic species formerly assigned to Pseudobolivina and Bigenerina, in: Proceedings of the Eighth International Workshop on Agglutinated Foraminifera, edited by: Kaminski, M. A. and Filipescu, S., Grzybowski Foundation Special Publications, 16, 19–27, 2011. 

Coleman, B. E.: Appendix 3: Foraminifera of the Oxford Clay and Kellaways Beds, in: The geology of the New Town of Milton Keynes, edited by: Horton, A., Coleman, B. E., Cox, B. M., Shephard-Thorn, E. R., Morter, A. A., Penn, I. E., Thurrell, R. G., and Ivimey-Cook, H. C., Institute of Geological Science Report, 74, 90–102, 1974. 

Cushman, J.: A Monograph of the Foraminifera of the North Pacific Ocean, Part 2, Textulariidae: Smithsonian Institute, Bulletin of the United States National Museum, 71, 1–108, 1911. 

Cushman, J. and Waters, J.: Additional Cisco Foraminifera from Texas, Contributions from the Cushman Laboratory for Foraminiferal Research, 4, 62–67, 1928. 

Cushman, J. A.: Ammodiscoides, a new genus of arenaceous foraminifera. Proceedings of the United States National Museum, Washington, DC, USA, 36, 423–425, 1909. 

Cushman, J. A.: New late Tertiary foraminifera from Vitilevu, Fiji: Contributions from the Cushman Laboratory for Foraminiferal Research, Sharon, Mass., USA, 7, 23–32, 1931. 

Cushman, J. A.: Upper Cretaceous Foraminifera of the Gulf Coastal region of the United States and adjacent areas, Geological Survey professional paper, 206, 1–241, 1946. 

Delage, Y. and Hérouard, E. J. É.: Traité de zoologie concrète: Vol.1, La Cellule et les Protozoaires, Paris, Schleicher Fréres, 107–155, 1896. 

d'Orbigny, A. D.: Mémoire sur les foraminiferes de la Craie Blanche du Basin de paris: Mémoires de la Société géologique de France, Paléontologie, Paris, France, 4, 51 pp., 1840. 

d'Orbigny, A. D.: Foraminiferes fossiles du Bassin Tertiaire de Vienne (Autriche), Gide, 2, 406 pp., 1846. 

Fox, L. R., Stukins, S., Hill, T., and Bailey, H.: New species of Cenozoic benthic foraminifera from the former British Petroleum micropalaeontology collection, J. Micropalaeontol., 37, 11–16,, 2018. 

Franke, A.: Die Foraminiferen der pommerschen Kreide: Greifswald, Univ., Geol.-Pal. Inst., Abh., Griefswald, Deutschland, 6, p. 69, 1925. 

Franke, A.: Die Foraminiferen und Ostracoden des Palaeocäns von Rugaard in Jütland und Sundkrogen bei Kopenhagen: Danmarks Geol. Unders., Kjobenhavn, Danmark, 2, 49 pp., 1927. 

Gauger, D.: Microfauna of the Hilliard Formation, in: Microfossils of the upper Cretaceous of northeastern Utah and southwestern Wyoming, eidted by: Peterson, R. H., Gauger, D. J., and Lankford, R. R., Bulletin of the Utah Geological and Mineralogical Survey, Salt Lake City, Utah, no. 47, 59–77, 1953. 

Gradstein, F. M., Kaminski, M., Berggren, W., Kristiansen, I., and d'Iorio, M.: Cenozoic biostratigraphy of the North Sea and Labrador shelf, Micropaleontology, 40, 152 pp., 1994. 

Heron-Allen, E. and Earland, A.: On the Recent and Fossil Foraminifera of the Shore-sands of Selsey Bill, Sussex – Part V., The Cretaceous Foraminifera, J. Microscopy, 30, 401–426, 1910. 

Hofker, J.: Les Foraminiferes du Crétacé supérieur do Contentin: 84 Congrès des Sociétés Savantes, Dijon, 368–397, 1959. 

Hofker Sr., J.: The Jurassic genus Reinholdella Brotzen, 1948 (Foraminifera), Paläontologische Zeitschrift, 26, 1–2, 1952. 

Jenkins, D. G. and Murray, J. W.: Stratigraphical Atlas of Fossil foraminifera, Ellis Horwood, 1989. 

Kaminski, M. A.: The year 2000 classification of the agglutinated foraminifera 2004, Grzybowski Foundation, in: Proceedings of the Sixth International Workshop on Agglutinated Foraminifera, edited by: Bubík, M. and Kaminski, M. A., Grzybowski Foundation Special Publication, 8, 237–255, 2004. 

Kaminski, M. A., Cetean, C. G., and Tyszka, J.: Nomenclature to describe the transition from multiserial to uniserial chamber arrangement in benthic foraminifera, J. Micropalaeontol., 30, 7–10,, 2011. 

King, C. and Hughes, M. J.:, Cainozoic micropalaeontological biostratigraphy of the North Sea, Report of the Institute of Geological Sciences, 82, 1–40, 1983. 

Kübler, J. and Zwingli, H.: Mikroskopische Bilder aus der Urwelt der Schweiz; Heft II Bürgersbibliothek Neujahrsblatt, Winterthur, 1–28, 1866. 

Kübler, J. and Zwingli, H.: Die Foraminiferen des Schweizerischen Jura, Winterthur, Schweiz, Steiner, 1–49, 1870. 

Lankester, E. R.: Protozoa, Encyclopaedia Britannica, 19, 830–866, 1885. 

Loeblich, A. R. and Tappan, H.: Some new and redefined genera and families of agglutinated foraminifera, Part I, J. Foramin. Res., 15, 91–104, 1985. 

Marie, P.: Foraminiferes de la Craie: Les foraminiferes de la Craie a Belemnitella mucronata du Bassin de Paris, Bulletin du Muséum National D'Histoire Naturelle, 12, p. 48, 1941. 

Marsson, T.: Die Foraminiferen der weissen Schreibkreide der Insel Rügen: Mittheilungen aus dem natur-wissenschaftlichen verein von Neu-Vorpommern und Rügen, Greifswald, Mitt., Berlin, Deutschland, 10, 115–196, 1878. 

Medd, A. W.: The Mesozoic stratigraphy of two boreholes near Worlaby, edited by: Richardson, G., Iremy-Cook, H., Medd, A., and Siddiqui, K., Humberside, HM Stationery Office, 1–24, 1979. 

Montagu, G.: Testacea Britannica, Supplement, Exeter, England, Printed by S. Woolmer, 1–183, 1808. 

Morris, P. H. and Coleman, B. E.: The Aalenian to Callovian, in: Stratigraphical Atlas of Fossil Foraminifera, edited by: Jenkins, D., G. and Murray, J. W., Chichester, UK, Ellis Horwood Ltd, 219–221, 1981.  

Parker, W. K. and Jones, T. R.: On the nomenclature of the foraminifera, Part II – On the species enumerated by Walker and Montagu, Annals and Magazine of Natural History, 4, 333–351, 1859. 

Reuss, A. E.: Ueber Ataxophragmium eine neue Foraminiferen-gattung aus der Familie der Uvellideen: Königl, Böhmischen Gesellschaft der Wissenschaften, 2, p. 52, 1860. 

Reuss, A. E.: Paläontologische Beiträge. Sitzungsberichte der Mathematisch-Naturwissenshaftlichen Classe der Kaiserlichen Akademie der Wissenschaften, Wien, 44, 304–342, 1862. 

Schijfsma, E.: The Foraminifera from the Hervian (Campanian) of Southern Limburg, Mededelingen van de Geologische Stichting, 7, 1–174, 1946. 

Schwager, C.: Quadro del proposto sistema di classificazione del foraminiferi con guscio, Bolletino Reale Comitato Geologico d'Italia, 8, 18–27, 1877. 

Sweicicki, A.: A foraminiferal biostratigraphy of the Campanian and Maastrichtian chalks of the United Kingdom, Ph.D., University of Plymouth, 66–67, 1980. 

Tairov, C. A.: New species of agglutinated foraminifera from Cretaceous deposits of north-eastern Azerbaijan, Transactions of the AzNII, 8, 221–244, 1959. 

Visser, A. M.: Monograph on the foraminifera of the type-locality of the Maestrichtian (South-Limburg, Netherlands), Leiden Geoligische Mededelingen, 16, 197–331, 1951. 

von Hagenow, F.: Monographie der Rügenschen Kreide- Versteinerunger, Abheilung 3: Mollusken: Neues Jarb. Min. Geogn. Geol. Petref.-Kunde, Stuttgart, Deutschland, p. 574, 1841. 

Short summary
This paper describes five new Mesozoic deep-water benthic foraminifera from the former British Petroleum microfossil reference collections at the Natural History Museum, London.